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Just for your interest: for some reason the value System\UnrealTorunament.ini->IpServer.UdpServerUplink->DoUplink=True changes automatically to False after starting the server. With setting DoUplink=True and then starting the server it connects to the given UdpServerUplink's, but with loading next map the uplink gets disabled (as seen in the logs).
I solved this by adding the parameter DoUplink=True for the UdpServerUplink, for example: Furthermore the definition of more than 3 UdpServerUplink's causes a SIGSEGV-crash as stated in 333networks-wiki.

Server details:

Spoiler

OS: Ubuntu 14.04.3 LTS
UT: server version 451
UT Framework: abfackelns Server Utilities (ASU) v0.6
UT mods: nexgen112, BarbiesWorld, MapvoteLA13, ServerAdds v1.04, SBMutatorFastWarShell-Rev2

update you Linux package ,as in what the wikia conserns you canot edit more then 3 lines of text in that section.
Otherwise you can get that fault on the server , you did described.
It just a limit of your Linux package or maby the ut 99 server files.

If you have more than 3 lines, your server may crash or not even start properly. Use trial-and-error to determine whether this is the case with your server.

best regardz:rjmno1